Research on Asian Rhinoceros

1965 Description of a New Subspecies of Rhinoceros, from Borneo. Saugetierkundl. Mitt. 13:128-131.

Description of Dicerorhinus sumatrensis harrissoni .

1967 The Rhinoceroses of Southeast Asia. Saugetierkundl. Mitt. 15:221-237.

1971 Species Characters in Rhinoceros Horns. Z.f. Saugetierk. 36:238-252.

1972 (with F. Kurt) Dicerorhinus Sumatrensis. Mammalian Species No. 21:1-6.

1980 (with C. Guérin). Le Rhinoceros sondaicus annamiticus (Mammalia, Perissodactyla) d'Indochine: distinction taxinomique et anatomique; relations phylétiques. Géobios 13:199-208.

1982 Asian Rhinoceroses - down but not out. Malayan Naturalist 36:11-22.

1982 The skulls of Asian Rhinoceroses: wild and captive. Zoo Biology 1:251-261.

1983 Phylogeny of the living species of Rhinoceros. Z. Zool. Syst. u. Evolutionsforschung 21:293-313.

1983 (with S. Chakraborty). The Calcutta collection of Asian Rhinoceros. Rec. Zool. Surv. India 80:251-263.

1984 (with W.A. Laurie and E.M. Lang) Rhinoceros unicornis. Mammalian species No. 211:1-6.

1993 Testing rhinoceros subspecies by multivariate analysis. Pp. 92-100 in O.A. Ryder (ed.) Rhinoceros Biology and Conservation. Zoological society of San Diego.

1999 Die Nashörner - Stammesgeschichte und Verwandtschaft. In Anonymous (ed.), Die Nashörner: Begegnung mit urzeitlichen Kolossen, 14-32. Furth: Filander Verlag.

Young female Bornean Rhinoceros (Dicerorhinus sumatensis harrissoni