Research on African Rhinoceros

1967 Geographic Variation in the Black Rhinoceros. Z.f. Saugetierk. 32:267-276.

1971 Species Characters in Rhinoceros Horns. Z.f. Saugetierk. 36:238-252.

1972 Ceratotherium Simum. Mammalian Species No. 8:1-6.

1975 Taxonomic notes on the White Rhinoceros Ceratotherium simum (Burchell, 1817). Saugetierkundl. Mitt. 23:200-212.

1978 The extinct Cape Rhinoceros, Diceros bicornis bicornis (Linnaeus, 1758). Saugetierkundl. Mitt. 26:117-28.

1983 Phylogeny of the living species of Rhinoceros. Z. Zool. Syst. u. Evolutionsforschung 21:293-313.

1993 Testing rhinoceros subspecies by multivariate analysis. Pp. 92-100 in O.A. Ryder (ed.) Rhinoceros Biology and Conservation. Zoological society of San Diego.

1994 (with A.K.Kes Hillman-Smith). Diceros bicornis. Mammalian Species, no.455, 8pp.

1999 Die Nashörner - Stammesgeschichte und Verwandtschaft. In Anonymous (ed.), Die Nashörner: Begegnung mit urzeitlichen Kolossen, 14-32. Furth: Filander Verlag.

Somali Black Rhinoceros (Diceros bicornis brucii) in Naples Zoo, about 1963

Northern White Rhinoceros (Ceratotherium simum cottoni), Dvur Kralove Zoo, about 1970

 

Conservation:

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